| 1 |
Stamens long exerted; corolla ca. 0.8 cm |
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35 Dracocephalum stamineum |
| + |
Stamens included or only short exserted; corolla 1-4.2 cm. |
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(2) |
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| 2 (1) |
Dwarf cespitose plants, stems 3-15 cm; branches numerous, crowded; leaves 3-5 mm, circular-ovate, pinnati- or palmatipartite. |
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(3) |
| + |
Stems erect or suberect, taller, unbranched or few branched; leaves more than 10 mm, simple to pinnatisect. |
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(5) |
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| 3 (2) |
Middle tooth of upper lip of calyx slightly wider than lateral teeth, upper lip teeth the same shape; leaves palmately 5-parted |
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30 Dracocephalum palmatoides |
| + |
Middle tooth of upper lip of calyx 2.5-5 × wider than lateral teeth, broadly ovate, lateral teeth lanceolate to narrowly triangular; leaves pinnatifid. |
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(4) |
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| 4 (3) |
Middle tooth of upper lip of calyx ca. 2.5 × wider than lateral teeth |
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31 Dracocephalum origanoides |
| + |
Middle tooth of upper lip of calyx 4-5 × wider than lateral teeth |
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32 Dracocephalum paulsenii |
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| 5 (2) |
Leaves pinnatisect, segments linear. |
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(6) |
| + |
Leaves undivided. |
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(13) |
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| 6 (5) |
Calyx conspicuously 2-lipped; leaves sometimes 2-pinnatifid |
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8 Dracocephalum bipinnatum |
| + |
Calyx of 5 subequal teeth, lanceolate to narrowly triangular; leaves clearly pinnatisect. |
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(7) |
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| 7 (6) |
Leaf sections 2- or 3-paired, crowded toward blade base, appearing palmatisect. |
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(8) |
| + |
Leaf segments not crowded at blade base. |
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(10) |
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| 8 (7) |
Flowers 2.5-2.8 cm; leaf segments suberect |
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5 Dracocephalum forrestii |
| + |
Flowers 3.5-4.2 cm; leaf segments oblique or subpatent. |
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(9) |
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| 9 (8) |
Calyx densely lanate-villous, teeth 1/4-1/3 as long as tube, teeth lanceolate-triangular, subulate- acuminate |
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6 Dracocephalum isabellae |
| + |
Calyx slightly scabrid, teeth ca. as long as tube, lanceolate, acute |
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7 Dracocephalum hookeri |
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| 10 (7) |
Corolla less than 1.5 cm |
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3 Dracocephalum propinquum |
| + |
Corolla more than 2 cm. |
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(11) |
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| 11 (10) |
Internodes 1-1.2 cm; leaves abaxially strigose on nerves |
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1 Dracocephalum taliense |
| + |
Internodes 2-6 cm; leaves abaxially densely white pubescent. |
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(12) |
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| 12 (11) |
Upper floral leaves much shorter than calyx |
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4 Dracocephalum tanguticum |
| + |
Upper floral leaves as long as to longer than calyx |
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2 Dracocephalum calophyllum |
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| 13 (5) |
Leaves entire. |
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(14) |
| + |
Leaves serrate or dentate. |
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(17) |
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| 14 (13) |
Bracts 2- or 3-serrulate; corolla 1.4-1.8 cm; teeth of upper lip of calyx divided almost to base, middle tooth of upper lip ca. 2 × as wide as lateral teeth; anthers glabrous. |
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(15) |
| + |
Bracts entire; corolla more than 1.8 cm; teeth of upper lip of calyx divided to nearly 1/2 its length, middle tooth of upper lip slightly wider than lateral teeth; anthers hairy. |
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(16) |
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| 15 (14) |
Leaves lanceolate to ovate-lanceolate, 1.5-3 × 4-8 mm, entire |
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15 Dracocephalum integrifolium |
| + |
Leaves linear, 5-7 × 0.8-1.2 mm, 2- or 3-serrulate; teeth with apical spines to 3 mm |
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14 Dracocephalum fruticulosum |
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| 16 (14) |
Stems subglabrous; calyx minutely hairy toward base, subglabrous toward apex; corolla 3.3-4 cm |
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34 Dracocephalum argunense |
| + |
Stems sparsely minute hairy toward base; calyx minutely hairy throughout; corolla 1.7-2.4 cm |
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33 Dracocephalum ruyschiana |
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| 17 (13) |
Calyx conspicuously 2-lipped. |
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(18) |
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Calyx inconspicuously 2-lipped. |
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(21) |
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| 18 (17) |
Calyx 2-lipped nearly to base |
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11 Dracocephalum breviflorum |
| + |
Calyx 2-lipped to ca. 1/2 its length. |
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(19) |
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| 19 (18) |
Leaves ovate, base ± cordate, crenate; bracts 3-8-spinescent-serrate; corolla white |
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9 Dracocephalum heterophyllum |
| + |
Leaves lanceolate or narrowly oblong, base cuneate to rounded, denticulate; bracts 1-3-spinescent-serrate; corolla blue-purple. |
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(20) |
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| 20 (19) |
Perennial herbs; leaves ± leathery, base cuneate, few spinescent-denticulate |
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10 Dracocephalum peregrinum |
| + |
Annual herbs; leaves herbaceous, base rounded to broadly cuneate, densely dentate with teeth not spinescent or sometimes basal teeth spinescent |
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12 Dracocephalum moldavica |
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| 21 (17) |
Middle tooth of upper lip ± same shape as lateral teeth, nearly as wide as to wider than lateral teeth. |
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(22) |
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Middle tooth of upper lip of calyx wider than lateral teeth, circular-ovate; lateral teeth broadly lanceolate to narrowly triangular. |
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(25) |
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| 22 (21) |
Basal leaves absent; cauline leaves many paired, short petiolate, elliptic to ovate-elliptic, 5-6 mm, 1-3-denticulate; calyx teeth 5, subequal; bracts spinescent-serrate |
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13 Dracocephalum psammophilum |
| + |
Basal leaves present. |
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(23) |
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| 23 (22) |
Cauline leaves many paired, broadly ovate; verticillasters in long spikes |
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16 Dracocephalum nutans |
| + |
Cauline leaves 2-6-paired, ovate to triangular-ovate; verticillasters crowded in capitula. |
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(24) |
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| 24 (23) |
Calyx minutely papillate hirtellous, glandular |
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19 Dracocephalum rigidulum |
| + |
Calyx pubescent to silky-villous, margin white ciliate |
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20 Dracocephalum imberbe |
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| 25 (21) |
Basal petiole much shorter than blade; blade ovate to broadly ovate, 0.6-2 cm, incised-dentate. |
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(26) |
| + |
Basal petiole slightly shorter to longer than blade. |
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(27) |
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| 26 (25) |
Corolla yellowish white, middle lobe of lower lip not spotted |
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17 Dracocephalum nodulosum |
| + |
Corolla purple, middle lobe of lower lip purple spotted |
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18 Dracocephalum hoboksarensis |
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| 27 (25) |
Corolla slightly exserted |
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22 Dracocephalum velutinum |
| + |
Corolla 1.5-2 × as long as calyx. |
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(28) |
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| 28 (27) |
Middle tooth of upper lip of calyx subtruncate. |
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(29) |
| + |
Middle tooth of upper lip of calyx rounded. |
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(30) |
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| 29 (28) |
Middle tooth of upper lip of calyx broadly elliptic, obtuse, short spinescent |
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26 Dracocephalum purdomii |
| + |
Middle tooth of upper lip of calyx trapeziform, truncate, denticulate |
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27 Dracocephalum truncatum |
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| 30 (28) |
Leaves abaxially densely white pubescent |
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21 Dracocephalum wallichii |
| + |
Leaves abaxially sparsely strigose. |
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(31) |
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| 31 (30) |
Petioles of middle cauline leaves 2-6 cm |
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29 Dracocephalum rupestre |
| + |
Petioles of middle cauline leaves less than 1.2 cm or obsolete. |
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(32) |
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| 32 (31) |
Floral leaves coarsely dentate; corolla 3.3-4 cm |
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28 Dracocephalum grandiflorum |
| + |
Floral leaves not coarsely dentate; corolla 2-3.5 cm. |
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(33) |
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| 33 (32) |
Leaves abaxially purplish, sparsely pubescent on veins or glabrous |
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25 Dracocephalum bullatum |
| + |
Leaves abaxially densely puberulent, sparsely floccose-villous on veins. |
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(34) |
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| 34 (33) |
Bracts subcircular, subentire |
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24 Dracocephalum imbricatum |
| + |
Bracts obovate to rhombic obovate, margin densely ciliate, 1-3-spinescent-serrate |
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23 Dracocephalum microflorum |
